Finally surgically transformed into a "pretty," sixteen-year-old Tally, now gorgeous and programmed to think only happy thoughts, is plagued by tangled memories of living in the Smoke, a rebel colony of "ugly" runaways hiding from the Special Circumstances authorities.
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Pretties picks up where Uglies left off. Tally has been turned pretty and dull-witted. Tally loves the pretty life of partying, drinking, and doing whatever she wants. She has a lot of fun with it, until one day at a costume party, she sees someone in a specials costume, and she starts feeling like something's not right with her world. Together with Zane, another pretty, Tally must track down the clue to her past as an ugly. When she does though, the repercussions of her choices afterward will continue long into the book. As Tally remembers the truth, she tries to figure out a way to help the other pretties and to save herself and Zane as well.
Tally is now a Perfect, and the only worries that occupy her brain is remaining "bubbly" and getting into the most popular clique in town. Until a letter written by herself in her ugly days turns her world upside down and reveals the horror of the transformation surgery.
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